Motility and self-organization of gliding <i>Chlamydomonas</i> populations

A population of gliding $C\phantom{\rule{0}{0ex}}h\phantom{\rule{0}{0ex}}l\phantom{\rule{0}{0ex}}a\phantom{\rule{0}{0ex}}m\phantom{\rule{0}{0ex}}y\phantom{\rule{0}{0ex}}d\phantom{\rule{0}{0ex}}o\phantom{\rule{0}{0ex}}m\phantom{\rule{0}{0ex}}o\phantom{\rule{0}{0ex}}n\phantom{\rule{0}{0ex}}a\phantom{\rule{0}{0ex}}s$ $r\phantom{\rule{0}{0ex}}e\phantom{\rule{0}{0ex}}i\phantom{\rule{0}{0ex}}n\phantom{\rule{0}{0ex}}h\phantom{\rule{0}{0ex}}a\phantom{\rule{0}{0ex}}r\phantom{\rule{0}{0ex}}d\phantom{\rule{0}{0ex}}t\phantom{\rule{0}{0ex}}i\phantom{\rule{0}{0ex}}i$ cells exhibit cluster formation. Cell motility alone in simulations is unable to reproduce the phenomenon, which can be resolved with inclusion mechanosensing.